Thanks for the Salmon Info Dave. That was much appreciated! Now lets get back to Steelhead!
How does a summer-run survive all that time in fresh water? They must go in a hypnotic state "trance" like sturgeon do in the winter.
Oh ya, I forgot that I killed one with a bullhead in its stomach!
Pacific salmon are the same genus as steelhead, and if the sculpin you found in a steelhead was even partially digested that would be relevant. Was it?
OK, let's try a different tact. If upstream migrating steelhead are able to digest food that means stomach excretions and enzymes are active … that also means an empty stomach would trigger a feeling of hunger and a search for food. Consider the Vedder gets a few thousand steelhead annually, average weight about 12 lbs; what do you think these fish would survive on if they had to rely on catching food?
If steelhead were indeed actively feeding and digesting what they captured, none would make it above the gauntlet of anglers and these fish would be long extinct. Being able to survive upstream migration without eating is a tactic that has worked for a long time for anadromous species.
Sure, it's hard to believe these fish survive for months without added nutrients, and may be a reason return spawners are very low in numbers, but it's true. It's much like how does a Stuart sockeye swim all that way up the Fraser simply by surviving on fat reserves?