In respect to enumerating adult Pacific Salmon species like Sockeye, there can be sex ratio differences annually, between geographic areas and run-timing groups. Some areas can have close to 50:50 while others can be skewed 60:40 (males or females being more prominent) for males or females. It is challenging to obtain a unbiased, representative sample to reflect the true sex ratio. If surveys have poor spatial (space) and temporal (time) representation that could bias the sex ratio. Ideally, ground surveys should be done at equal frequency intervals over time and cover the extent of spawning for a particular stream. Seems like an easy thing to satisfy, but in reality it can be challenging. Fish behaviour, predators and environmental conditions are factors that play into this.
For instance, males typically die sooner than females and leave the system making them less recoverable for survey crews. Secondly, extreme environmental conditions (water temperatures or water levels) can impact sex ratio. For instance, female Sockeye can experience higher enroute mortality if migratory conditions in the Fraser are extreme. Thirdly, predators (specifically bears) on spawning grounds can remove salmon carcasses making them less available for crews to recover in order to determine sex ratio. If salmon abundances on a particular stream are low and there many bears are feeding during the day then it may not leave much for crews to count in order to determine sex ratio - especially if they are only visiting the stream once every 7 days. This may necessitate survey frequency to be even more often.